遗迹化石<i>Zoophycos</i>的系统修订与分类

张立军; 龚一鸣

doi:10.3799/dqkx.2012.S1.007

遗迹化石Zoophycos的系统修订与分类

doi: 10.3799/dqkx.2012.S1.007

张立军^{1, 2,},
龚一鸣^1, ,

1.
中国地质大学生物地质与环境地质国家重点实验室，湖北武汉 430074
2.
河南理工大学资源环境学院，河南省生物遗迹与沉积矿产河南省重点实验室，河南焦作 454003

基金项目:

国家自然科学基金 40872001

国家自然科学基金 41072252

国家自然科学基金 40921062

国家重点基础研究计划“973”项目 2011CB808800

河南省生物遗迹与成矿过程重点实验室开放基金 OTMP1001

详细信息

作者简介:
张立军(1982-)，男，博士研究生，主要从事遗迹化石与地球生物学研究.E-mail: Ljzhanghpu@gmail.com

通讯作者:
龚一鸣，E-mail: ymgong@cug.edu.cn

中图分类号: P52
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出版历程
- 收稿日期: 2011-07-26
- 网络出版日期: 2021-11-15
- 刊出日期: 2012-05-01

Systematic Revision and Ichnotaxonomy of Zoophycos

ZHANG Li-jun^{1, 2
,},
GONG Yi-ming^{1
, ,}

1.
State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Wuhan 430074, China
2.
Institute of Resources and Environment, Key Laboratory of Biogenic Traces & Sedimentary Minerals of Henan Provinces, Henan Polytechnic University, Jiaozuo 454003, China

摘要

摘要: 三维形态结构复杂、分布广泛的Zoophycos一直是众多遗迹学家研究的热点.通过对1855年至2011年发表的具有详细形态描述并配有图版的动藻迹(Zoophycos)文献进行系统整理, 基于详实的Zoophycos形态结构特征, 系统修订了Zoophycos属, 将Zoophycos主要划分为9个种, 即Zoopycos cauda-galli Vanuxem, 1842；Zoophycos curtain Vanuxem, 1842；Zoophycos velum Vanuxem, 1842；Zoophycos brianteus Massalongo, 1855；Zoophycos villae Massalongo, 1855；Zoophycos liasinus Fischer-Ooster, 1858；Zoophycos flabelliformis Fischer-Ooster, 1858；Zoophycos insignis Squinabol, 1890和Zoophycos rhodensis Bromley and Hanken, 2003; 将Spirophyton主要划分为2个种：Spirophyton typum Hall, 1863和Spirophyton eifeliense Kayser, 1872.将Zoophycos与Spirophyton共同归入Zoophycos Group.
- 动藻迹 /
- 螺藻迹 /
- 遗迹化石 /
- 属种修订 /
- 地球生物学 /
- 形态学
Abstract: Zoophycos is one of the most widespread but still enigmatic trace fossils, especially with regard to morphology and ethology surrounding its 3-D helically coiled spreite structures. The genus Zoophycos is systematically revised based on the systematic researches of the literature with detailed description and plate from 1855 to 2011. The ichnogenus Zoophycos is revised into 9 ichnospecies, which include Zoophycos cauda-galli Vanuxem, 1842; Zoophycos curtain Vanuxem, 1842; Zoophycos velum Vanuxem, 1842; Zoophycos brianteus Massalongo, 1855; Zoophycos villae Massalongo, 1855; Zoophycos liasinus Fischer-Ooster, 1858; Zoophycos flabelliformis Fischer-Ooster, 1858; Zoophycos insignis Squinabol, 1890; Zoophycos rhodensis Bromley and Hanken, 2003. The ichnogenus Spirophyton is composed of Spirophyton typum Hall, 1863 and Spirophyton eifeliense Kayser, 1872. The Zoophycos and Spirophyton are grouped into Zoophycos Group.
- Zoophycos /
- Spirophyton /
- trace fossils /
- revision of genera and species /
- geobiology /
- morphology

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图 1 Zoophycos brianteus、Zoophycos cauda-galli和Fucoides circinatus形态结构特征

a.Zoophycos brianteus Massalongo, 1855原始素描图(转引自Olivero, 2007); b.意大利维罗纳博物馆Zoophycos brianteus Massalongo, 1855补选模式种标本(编号ICN3)(左侧)与原始素描图(右侧)(转引自Olivero, 2007); c.意大利维罗纳博物馆Zoophycos brianteus Massalongo, 1855副模标本(转引自Olivero, 2007); d~e.美国纽约州泥盆纪钙质砂岩水平面中Zoophycos cauda-galli素描图(Vanuxem, 1842), f.Fucoides circinatus(Brongniart, 1828); g.瑞典Husaby地区Bishop城堡废墟石墙上的Fucoides circinatus(Jensen and Bergström, 1995)

Fig. 1. Morphological characteristics of Zoophycos brianteus、Zoophycos cauda-galli and Fucoides circinatus

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图 2 修订后世界各地保存较好的Zoophycos brianteus形态

a~c.Zoophycos circinnatus, Pli ka, 1968; d.Zoophycos, Webby, 1969; e.Spirophyton crassum Hall, 1863; f.意大利维罗纳博物馆Zoophycos brianteus Massalongo, 1855副模标本(转引自Olivero, 2007); g.Zoophycos circinnatus, Pli ka, 1969; h.Zoophycos, Olivero, 2003; i.Zoophycos circinnatus, Pli ka, 1969; j~k.Spirophyton Type ‘A’, Plumstead, 1967

Fig. 2. Representative samples of revised Zoophycos brianteus occurred in the world

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图 3 Zoophycos curtain和Zoophycos velum素描(Vanuxem, 1842)

Fig. 3. Original drawing of Zoophycos curtain and Zoophycos velum

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图 4 Zoophycos villae形态结构特征

a.Zoophycos villae Massalongo, 1855原始素描图(转引自Olivero, 2007); b.意大利维罗纳博物馆Zoophycos villae Massalongo, 1855副模标本(转引自Olivero, 2007); c.Zoophycos villae Massalongo, 1855补选模式种标本(左侧)(编号ICN2)与原始素描图(右侧)(转引自Olivero, 2007); d.Zoophycos villae Massalongo, 1855补选模式种标本(侧面观察)(编号ICN2)与其素描简图(转引自Olivero, 2007)

Fig. 4. Morphological characteristics of Zoophycos villae

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图 5 修订后的Zoophycos flabelliformis的形态结构特征

a.Taonurus flabelliformis Fischer-Oster, 1858; b.Zoophycos, Miller and D'Alberto, 2001; c~d.Spirophyton Type ‘B’, Plumstead, 1967; e.Zoophycos, Miller and D'Alberto, 2001

Fig. 5. The illustration showing the representative Zoophycos flabelliformis revised

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图 6 修订后的Zoophycos liasinus的形态结构特征

Fig. 6. The illustration showing the representative Zoophycos liasinus revised

a.Taonurus liasinus Fischer-Ooster, 1858; b.Zoophycos ferrum equinum Heer, 1865; c.Zoophycos scoparius Thiol, 1865; d~e.Zoophycos Type D, Patel et al., 2009

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图 7 修订后Zoophycos insignis的形态结构特征

a.Zoophycos insignis, Barsanti, 1902; b.Zoophycos insignis, Uchman, 1999

Fig. 7. The illustration showing the representative Zoophycos insignis revised

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图 8 修订后Zoophycos rhodensis的主要形态

a.Zoophycos rhodensis Bromley and Hanken, 2003; b.Zoophycos, Lewis, 1970

Fig. 8. The illustration showing the representative Zoophycos rhodensis revised

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图 9 修订后Spirophyton typum和Spirophyton eifeliense形态

a.俯视素描图Spirophyton typum Hall, 1863; b.俯视素描图Spirophyton typum Hall, 1863; c.三维形态恢复图Spirophyton typum Hall, 1863; d.俯视图Spirophyton eifeliense Kayser, 1872

Fig. 9. The illustration showing the representative Spirophyton typum and Spirophyton eifeliense revised

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图 10 Zoophycos疑问种

a~b.Zoophycos brianteus Fisher-Ooster, 1858; c~d.Cancellophycus marioni Saporta, 1887; e~h.Zoophycos taiyuanensis, 杨式溥, 2004; i~j.Zoophycos xuzhouensis Long, 1987(转杨式溥等，2004); k.Zoophycos circinnatus(转杨式溥等，2004); l.Zoophycos luchengensis(转杨式溥等, 2004); m~n.Zoophycos wudaensis Yang, 1984(转杨式溥等, 2004)

Fig. 10. Questionable Zoophycos species

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图 11 Zoophycos与Spirophyton对比

Zoophycos 3D形态据Löwemark and Werner, 2001修改; Spirophyton 3D形态据Hall, 1863修改

Fig. 11. Sketch showing the differences between Zoophycos and Spirophyton

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图 12 Rhizocorallium-like Zoophycos与Rhizocorallium对比

Zoophycos 3D形态据Bromley et al., 1999修改; Rhizocorallium 3D形态据Fürsich, 1974修改

Fig. 12. Sketch showing the differences between Rhizocorallium-like Zoophycos and Rhizocorallium

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图 13 Zoophycos Group谱系图

Fig. 13. The pedigree tree of Zoophycos Group

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表 1 设想的Zoophycos和Spirophyton特征

Table 1. Assumed morphological codes of Zoophycos and Spirophyton

代码	数字	特征
(1)柱形管	0	无柱形管
(1)柱形管	1	有柱形管
(2)边缘管	0	无边缘管
(2)边缘管	1	有边缘管
(3)大蹼纹	0	无大蹼纹
(3)大蹼纹	1	有大蹼纹
(4)小蹼纹	0	无小蹼纹
(4)小蹼纹	1	有小蹼纹
(5)水平面上形态结构	0	圆形蹼层
	1	椭圆形蹼层
	2	U形蹼层
	3	U形且底端转折或扩散形蹼层
	4	漏斗形蹼层
	5	螺旋桨形蹼层
	6	花瓣形蹼层
	7	裙带形蹼层
	8	J形蹼层
(6)纵切面柱形管特征	0	纵切面柱形管呈J形
(6)纵切面柱形管特征	1	纵切面柱形管呈U形

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表 2 Zoophycos和Spirophyton数字矩阵

Table 2. The digital matrix of Zoophycos and Spirophyton

	(1)	(2)	(3)	(4)	(5)	(6)
Z.caudagalli	1	1	1	1	1	0
Z.brianteus	1	0	1	1	0	0
Z.villae	0	1	1	1	8	0
Z.flabelliformis	1	1	1	0	6	1
Z.liasinus	0	1	1	0	2	1
Z.insignis	1	1	1	1	5	1
Z.rhodensis	1	1	1	1	7	1
Z.curtainensis	1	1	1	0	3	1
Z.velum	1	1	1	0	4	0
S.typum	1	1	1	0	0	0
S.eifeliense	1	1	1	0	0	0

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参考文献(93)

Antun, P., 1950. Sur les Spirophyton de l'Emsien de l'Oesling (Grand-Duché de Luxembourg). Annales de Ia Société GéoIogique de Belgique, 73: 241-261.
Barsanti, D.L., 1902. Considerazioni sorpa IL Genere Zoophycos. Atti Societa Toscana Scienze Naturrali, 18: 68-95.
Bellotti, P., Valeri, P., 1976. Zoophycos burrows in the Ammonitico Rosso Superiore Formation of Pellecchia Mount (Lucretili Mounts). Bollettino della Societa Geologica Italiana, 97: 21-34.
Bellotti, P., Valeri, P., 1978. L'Influenza Dell'Ambiente sedimentario sull'assetto elicoidale delle strutture a Zoophycos. Bollettino della Societa Geologica Italiana, 97: 675-685.
Bellotti, P., Valeri, P., 1987. New data on the ecology of Zoophycos in pelagic sediments of Garulla, Sibillini Mountains, Marches Apennines. Bollettino della Societa Geologica Italiana, 106: 813-817.
Bischoff, B., 1968. Zoophycos, a polychaete annelid, Eocene of Greece. Journal of Paleontology, 42(6): 1439-1443. doi: 10.2307/1302293
Bradley, J., 1973. Zoophycos and Umbellula (Pennatulacea): their synthesis and identity. Palaeogeography, Palaeoclimatology, Palaeoecology, 13(2): 103-128. doi: 10.1016/0031-0182(73)90039-4
Bromley, R.G., Ekdale, A.A., Asgaard, U., 1999. Zoophycos in the Upper Cretaceous chalk of Denmark and Sweden. Greifwalder Geowissenscha ftliche Beirtage, 6: 133-142.
Bromley, R.G., Hanken, N.M., 2003. Structure and function of large, lobed Zoophycos, Pliocene of Rhodes, Greece. Palaeogeography, Palaeoclimatology, Palaeoecology, 192(1): 79-100. doi: 10.1016/S0031-0182(02)00680-6
Brongniart, A., 1828. Histoire des végétaux fossiles ou recherches botaniques et géologiques. Tome premier, Paris, 82-84.
Carvalho, C.N.D., Rodrigues, N.P.C., 2003. Los Zoophycos del Bajociense-Bathoniense de la Praia da Mareta (Algarve, Portugal): arquitectura y finalidades en régimen de dominancia ecológica. Revista Española de Paleontologia, 18(2): 229-241.
Cave, M.P., 1982. Occurrence of Zoophycos in Oretian rocks of the Murihiku Supergroup, South Otago, New Zealand. New Zealand Journal of Geology and Geophysics, 25: 367-369. doi: 10.1080/00288306.1982.10421501
Ekdale, A.A., Lewis, D.W., 1991. The New Zealand Zoophycos revisited: morphology, ethology, and paleoecology. Ichnos, 1(3): 183-194. doi: 10.1080/10420949109386351
Fischer-Ooster, C.V., 1858. Die Fossilen Fucoiden der Schweizer-Alpen, nebst Eröterungen über deren geologisches Alter. Huber, Bern, 42-44.
Fu, S.P., Werner, F., 1995. Is Zoophycos a feeding trace?N. Jb. Geol. Paläont. Abh., 195(1-3): 37-47. doi: 10.1127/njgpa/195/1995/37
Fürsich, F.T., 1974. Ichnogenus Rhizocorallium. Paläont. Z., 48(1-2): 16-28. doi: 10.1007/BF02986987
Fraaye, R.H.B., Werver, O.P., 1990. Trace fossils and their environmental significance in Dinatian carbonates of Belgium. Paläont. Z., 64(3-4): 367-377. doi: 10.1007/BF02985726
Gaillard, C., Olivero, D., 1993. A new paleoecologic interpretation of Zoophycos Massalongo, 1855. C.R. Acad. Sci. Pairs, 316(11): 823-830.
Gaillard, C., Hennebert, M., Olivero, D., 1999. Lower Carboniferous Zoophycos from the Tournai area (Belgium): environmental and ethologic significance. Geobios, 32(4): 513-524. doi: 10.1016/S0016-6995(99)80001-1
Gaillard, C., Racheboeuf, P.R., 2006. Trace fossils from nearshore to offshore environments: Lower Devonian of Bolivia. Journal of Paleontology, 80(6): 1205-1226. doi: 10.1666/0022-3360(2006)802.0.CO;2
Goldring, R., Taylor, A.M., Hughes, G.W., 2005. The application of ichnofabrics towards bridging the dichotomy between siliciclastic and carbonate shelf facies: examples from the Upper Jurassic Fulmar Formation (UK) and the Jubaila Formation (Saudi Arabia). Proceedings of the Geologists' Association, 116(3-4): 235-249. doi: 10.1016/S0016-7878(05)80044-6
Gong, Y.M., Hu, B., Qi, Y.A., et al., 2010. Progress in Ichnofabric and Ichnocoenosis. China Coal Industry Publishing House, Beijing (in Chinese).
Hall, J., 1863. Observations upon some spiral growing fucoidal remains of the Paleozoic rocks of New York. New York Cabinet, 16th Annual Report, 76-83.
Häntzschel, W., 1962. Trace fossils and problematica. In: Moore, R.C. ed., Treatise on Invertebrate Paleontology, part W, Miscellanea: condonts, conoidal shells of uncertain affinities, worms, trace fossils and Problematica. Geological Society of America, New York and University Kansas Press, Lawrence, Kansas, 177-245.
Häntzschel, W., 1975. Trace fossils and problematica. In: Teichert, C., ed., Treatise on invertebrate paleontology, part W, Miscellanea, Supplement 1. Geological Society of America, New York and University of Kansas Press, Lawrence, Kansas, 1-269.
Heer, O., 1865. Die Urwelt der Schweiz. Friedrich Schultheβ, Zürich, 140-142.
Jensen, S., Bergström, J., 1995. The trace fossil Fucoides circinatus Brongniart, 1828, from its type area, Västergötland, Sweden. GEF, 117(4): 207-210. doi: 10.1080/11035899509546217
Kansha, Y., 1974. Ichnofossil Zoophycos from the Arita Formation at Yuasa-cho, Wakayama Prefecture, Japan. Journal of Geological Society of Japan, 80(8): 381-382. doi: 10.1292/geosoc.80.381
Kayser, F.H.E., 1872. Neue Fossilien aus dem rheinischen Devon. Deutsch. Geol. Gesell., Zeitschr, 24: 691-700.
Kotake, N., 1989. Paleoecology of the Zoophycos producers. Lethaia, 22: 327-341. doi: 10.1111/j.1502-3931.1989.tb01349.x
Kotake, N., 1990. Mode of ingestion and egestion of the Chondrites and Zoophycos producers. Journal of Geological Society of Japan, 96(10): 859-868. doi: 10.1292/geosoc.96.859
Kotake, N., 1991. Non-selective surface deposit feeding by the Zoophycos producers. Lethaia, 24: 379-385. doi: 10.1111/j.1502-3931.1991.tb01489.x
Kotake, N., 1992. Deep-sea echiurans: possible producers of Zoophycos. Lethaia, 25: 311-316. doi: 10.1111/j.1502-3931.1992.tb01400.x
Kotake, N., 1993. Tiering of trace fossil assemblages in Plio-Pleistocene bathyal deposits of Boso Peninsula, Japan. Palaios, 8: 544-553. doi: 10.2307/3515031
Kotake, N., 1994. Population paleoecology of the Zoophycos-producing animal. Palaios, 9: 84-91. doi: 10.2307/3515081
Kues B.S., 2005. The trace fossil Zoophycos in the Sandia Formation of north-central New Mexico. New Mexico Geology, 27(3): 70-71.
Lewis, D.W., 1970. The New Zealand Zoophycos. New Zealand Journal of Geology and Geophysics, 13(2): 295-315. doi: 10.1080/00288306.1970.10423968
Long, Y.Z., Wang, H.W., He, K.L., 1987. The discover of trace fossils from Xuzhuang Formation of Middle Cambrian at Dabeiwang distinct, Xuzhou, Jiangsu and its paleoenvironmental significance. Lithofacies and paleogeography, 13(3): 17-29 (in Chinese).
Löwemark, L., Werner, F., 2001. Dating errors in high-resolution stratigraphy: a detailed X-ray radiograph and AMS-¹⁴C study of Zoophycos burrows. Marine Geology, 177(3-4): 191-198. doi: 10.1016/S0025-3227(01)00167-0
Massalongo, A., 1855. Zoophycos, novum genus Plantarum fossilium. Typis Antonellianis, Veronae, 45-52.
Marintsch, E.J., Finks, R.M., 1978. Zoophycos size may indicate environmental gradients. Lethaia, 11: 273-279. doi: 10.1111/j.1502-3931.1978.tb01880.x
Miller, W. Ⅲ, D'Alberto, L., 2001. Paleoethologic implications of Zoophycos from Late Cretaceous and Paleocene limestones of the Venetian Prealps, northeastern Italy. Palaeogeography, Palaeoclimatology, Palaeoecology, 166(1-2): 237-247. doi: 10.1016/S0031-0182(00)00211-X
Olivero, D., 1995. La trace fossile Zoophycos historique et interprétations actuelles. Bollettino del Museo Regionale di Scienze Naturali (Torino), 13(1): 5-34.
Olivero, D., 2003. Early Jurassic to Late Cretaceous evolution of Zoophycos in the French Subalpine basin (southeastern France). Palaeogeography, Palaeoclimatology, Palaeoecology, 192(1-4): 59-78. doi: 10.1016/S0031-0182(02)00679-X
Olivero, D., 2007. Zoophycos and the role of type specimens in ichnotaxonomy. In: Miller, W. Ⅲ, ed., Trace fossils concepts, problems, prospects. Elservier, Amesterdam, 219-231.
Olivero, D., Gaillard, C., 1996. Paleoecology of Jurassic Zoophycos from south-eastern France. Ichnos, 4: 249-260. doi: 10.1080/10420949609380135
Osgood, R.G., Szmuc, E.J., 1972. The trace fossil Zoophycos as an indicator of water depth. Bulletins of American Paleontology, 62(271): 5-22.
Pavlovec, R., Pav i, J., 1971. Zoophycos (Annelida, Polychaeta) in the strata of Podsabotin in western Slovenia. Geologija, 14: 63-66.
Patel, S.J., Desai, B.G., Shukla, R., 2009. Paleoecological significance of the trace fossils of Dhosa Oolite Member (Jumara Formation), Jhura dome, Mainland Kachchh, Western India. Journal Geological Society of India, 74: 601-614. doi: 10.1007/s12594-009-0175-9
Pli ka, M., 1965. Origin of fossil Zoophycos. Nature, 208(5010): 579. doi: 10.1038/208579a0
Pli ka, M., 1968. Zoophycos, and a proposed classification of sabellid worms. Journal of Paleontology, 42(3): 836-849.
Pli ka, M., 1969. Methods for the study of Zoophycos and similar fossils. New Zealand Journal of Geology and Geophysics, 12(2-3): 551-573. doi: 10.1080/00288306.1969.10420298
Plumstead, E.P., 1967. A general review of the Devonian fossil plants found in the cape system of South Africa. Palaeontologia Africana, 10: 1-83.
Reid, C.M., James, N.P., Beauchamp, B., et al., 2007. Faunal turnover and changing oceanography: Late Palaeozoic warm-to-cool water carbonates, Sverdrup basin, Canadian Arctic Archipelago. Palaeogeography, Palaeoclimatology, Palaeoecology, 249(1-2): 128-159. doi: 10.1016/j.palaeo.2007.01.007
Ride, W.D.L., Cogger, H.G., Dupuis, C., et al., 1999. International code of zoological nomenclature. The Natural History Museum, London, 1-124.
Seilacher, A., 1967. Fossil behavior. Scientific American, 217: 72-80. doi: 10.1038/scientificamerican0867-72
Squinabol, S., 1890. Alghe e pseudoalghe fossili italiane. Atti Società Linguistica Scienze Naturali Geografiche, 1(1-2): 29-49, 166-199.
Stevens, G.R., 1968. The Amuri fucoid. New Zealand Journal of Geology and Geophysics, 11(1): 253-261. doi: 10.1080/00288306.1968.10423693
Saporta, G., 1887. Nouveaux documents relativs aux organismes problematiques des anciennes mers. Bull. Soc. geol. France, 15(3): 286-302.
Tavernier, A., 1988. L'Ichnofacies a Zoophycos del <<Flysch DI Faeto>>(Appennino Dauno). Considerazioni paleoambientali. Memorie della Societa Geologica Italiana, 41: 605-608.
Uchman, A., 1995. Taxonomy and palaeoecology of flysch trace fossils: the Marnoso-arenacea Formation and associated facies (Miocene, Northern Apennines, Italy). Beringeria, 15: 3-115.
Uchman, A., 1999. Ichnology of the Rhenodanubian Flysch (Lower Cretaceous-Eocene) in Austria and Germany. Beringeria, 25: 67-173.
Vanuxem, L., 1842. Geology of New York part Ⅲ survery of the third geological district. White and Visscher, 1-306.
Villa, A., 1844. Memoria sulla costituzione geologica e geognostica della Brianza. Milano, 1-46.
Warne, J.E., Kennedy, W.J., Schneidermann, N., 1973. Biogenic sedimentary structures (trace fossils) in leg 15 cores. Deep Sea Drilling Project Intitial Reports, 15: 813-831.
Webby, B.D., 1969. Trace fossils Zoophycos and Chondrites from the Tertiary of New Zealand. New Zealand Journal of Geology and Geophysics, 12: 208-214. doi: 10.1080/00288306.1969.10420234
Wetzel, A., Werner, F., 1981. Morphology and ecological significance of Zoophycos in deep-sea sediments off NW Africa. Palaeogeography, Palaeoclimatology, Palaeoecology, 32: 185-212. doi: 10.1016/0031-0182(80)90040-1
Yang, S.P., 1984. The discovery of Zoophycus in China and its environmental significance. Oil & Gas Geology, 5(3): 228-235 (in Chinese with English abstract).
Yang, S.P., 1985. Advances in palaeoichnological research in China. Geological Review, 31(3): 197-203 (in Chinese with English abstract).
Yang, S.P., Lin, W.Q., Tang, D.Z., 1988. Trace fossils. In: Hou, H.F., et al., eds., Devonian strata, paleontology and facies of the Longmenshan Mountain region. Geological Publishing Press, Beijing, 356-363 (in Chinese).
Yang, S.P., 1990. Palaeoichnology. Geological Publishing Press, Beijing (in Chinese).
Yang, S.P., 1994. Cambrian-Ordovician trace fossils from Kuruktag Mt., North-eastern part of Tarim basin, Xinjiang. Geoscience, 8(4): 371-380 (in Chinese with English abstract).
Yang, S.P., Zhang, J.P., Yang, M.F., 2004. Trace fossils of China. Science Press, Beijing (in Chinese).
Yang, S.P., Zheng, Z.C., 1990. Trace fossils from Gongwusu Formation of Middle Ordovician in Wuhai, Neimenggu. Ningxia Geology, 2: 102-115 (in Chinese).
Zhang, L.J., Gong, Y.M., 2009. New discovery of Late Devonian plant and trace fossils in Hougaoping, Guangyuan, Sichuan. Journal of Stratigraphy, 33(2): 138-146 (in Chinese with English abstract).
Zhang, L.J., Gong, Y.M., 2011. Spatiotemporal distribution and controlling factor of the Late Paleozoic Zoophycos in South China. Science & Technology Review, 29(31): 18-28 (in Chinese with English abstract).
Zhang, X.P., 1991. Trace fossils from Tiaomajian Formation of Middle Devonian in Xinhua, Xiantan and other areas. Hunan Geology, 10(2): 97-104 (in Chinese with English abstract).
Zhang, X.P., Qing, S.K., 1988. Upper Devonian trace fossils from Xinhua County, Hunan and their sedimentary environment. Oil & Gas Geology, 9(3): 252-260 (in Chinese with English abstract).
Zhang, X.P., Qing, S.K., 1992. Trace fossils and sedimentary environments of Lower Carboniferous at Xinhua, Hunan. Acta Palaeontologica Sinica, 31(5): 605-617 (in Chinese with English abstract).
龚一鸣, 胡斌, 齐永安, 等, 2010. 遗迹组构和遗迹群落研究新进展. 北京: 煤炭工业出版社.
龙耀珍, 王宏伟, 何康林, 1987. 江苏徐州大北望地区中寒武统徐庄组遗迹化石的发现及其环境意义. 岩相古地理, 13(3): 17-29.
杨式溥, 张建平, 杨美芳, 2004. 中国遗迹化石. 北京: 科学出版社.
杨式溥, 郑昭昌, 1990. 内蒙古乌海市中奥陶统公乌素组遗迹化石. 宁夏地质, 2: 102-115.
杨式溥, 1984. Zoophycus在中国的发现及其环境意义. 石油与天然气地质, 5(3): 228-235.
杨式溥, 1985. 我国遗迹化石研究的新进展. 地质论评, 31(3): 197-203.
杨式溥, 林文球, 唐德章, 1988. 遗迹化石. 侯鸿飞, 等编, 四川龙门山地区泥盆纪地层古生物及沉积相. 北京: 地质出版社, 356-363.
杨式溥, 1990. 古遗迹学. 北京: 地质出版社.
杨式溥, 1994. 新疆塔里木盆地东北部库鲁克塔格地区寒武-奥陶纪遗迹化石. 现代地质, 8(4): 371-380.
张立军, 龚一鸣, 2009. 四川后高坪地层晚泥盆世植物和遗迹化石的新发现. 地层学杂志, 33(2): 138-146. doi: 10.3969/j.issn.0253-4959.2009.02.003
张立军, 龚一鸣, 2011. 华南晚古生代Zoophycos时空分布及其控制因素. 科技导报, 29(31): 18-28. doi: 10.3981/j.issn.1000-7857.2011.31.002
张欣平, 卿上康, 1988. 湖南新化上泥盆统的遗迹化石及其沉积环境. 石油与天然气地质, 9(3): 252-260.
张欣平, 卿上康, 1992. 湖南新化下石炭统遗迹化石及其沉积环境. 古生物学报, 31(5): 605-617.
张欣平, 1991. 新化、湘潭等地跳马涧组的遗迹化石. 湖南地质, 10(2): 97-104.