| Citation: | Mao Ningfeng, Chen Chen, Ye Qin, An Zhihui, Hu Jun, Tong Jinnan, 2025. Macroscopic Carbonaceous Compression Fossils from Late Ediacaran Miaohe Member at the Lianhua Section in Shennongjia Area. Earth Science, 50(9): 3704-3716. doi: 10.3799/dqkx.2025.107 | 
	                | 
					 An, Z. H., Jiang, G. Q., Tong, J. N., et al., 2015. Stratigraphic Position of the Ediacaran Miaohe Biota and Its Constrains on the Age of the Upper Doushantuo δ13C Anomaly in the Yangtze Gorges Area, South China. Precambrian Research, 271: 243-253.  https://doi.org/10.1016/j.precamres.2015.10.007 
						
					 | 
			
| 
					 An, Z. H., Ye, Q., Hu, J., et al., 2023. Stratigraphic Position of Cryogenian Songluo Biota in Shennongjia Area. Earth Science, 48(10): 3799-3811 (in Chinese with English abstract). 
						
					 | 
			
| 
					 An, Z. H., Ye, Q., Tong, J. N., et al., 2024. Stratigraphic Sequence of the Ediacaran Dengying Formation around the Huangling Anticline, Western Hubei Provence: Implications for Regional Stratigraphic Correlation. Earth Science, 49(8): 2992-3009 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Anderson, E. P., Schiffbauer, J. D., Xiao, S. H., 2011. Taphonomic Study of Ediacaran Organic-Walled Fossils Confirms the Importance of Clay Minerals and Pyrite in Burgess Shale-Type Preservation. Geology, 39(7): 643-646.  https://doi.org/10.1130/g31969.1 
						
					 | 
			
| 
					 Butterfield, N. J., 2007. Macroevolution and Macroecology through Deep Time. Palaeontology, 50(1): 41-55.  https://doi.org/10.1111/j.1475-4983.2006.00613.x 
						
					 | 
			
| 
					 Chen, M. E., Xiao, Z. Z., Yuan, X. L., 1994. A New Assemblage of Megafossils-Miaohe Biota from Upper Sinian Doushantuo Formation, Yangtze Gorges. Acta Palaeontologica Sinica, 33(4): 391-403 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Chen, X. H., Wang, X. F., 2002. Morphology and Systematics of Fossils in the Sinian Wulingshan Biota from Western Hunan Province. Geological Bulletin of China, 21(10): 638-645 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Chen, X. H., Wang, X. F., Wang, C. S., et al., 1999. A Preliminary Study on Carbonaceous Megafossils from the Late Sinian Liuchapo Formation of West Hunan. Geology and Mineral Resources of South China, 15(2): 15-30 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Croce, M. E., Gauna, M. C., Fernández, C., et al., 2021. Biology and Ecology of the Benthic Algae. In: Fiori, S. M., Pratolongo, P. D., eds., The Bahía Blanca Estuary. Springer International Publishing, Cham, 113-151.  
						
					 | 
			
| 
					 Ding, L. F., Li, Y., Hu, X. S., et al., 1996. Sinian Miaohe Biota. Geological Publishing House, Beijing, (in Chinese). 
						
					 | 
			
| 
					 Gu, H. D., Hu, J., An, Z. H., et al., 2021. Sedimentary Characteristics of Doushantuo Formation in Shennongjia Area: Implications for "West Hubei Trough". Earth Science, 46(8): 2958-2972 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Hu, J., Li, C., Tong, J. N., et al., 2020. Glacial Origin of the Cryogenian Nantuo Formation in Eastern Shennongjia Area (South China): Implications for Macroalgal Survival. Precambrian Research, 351: 105969.  https://doi.org/10.1016/j.precamres.2020.105969 
						
					 | 
			
| 
					 Hua, H., Cai, Y. P., Min, X., et al., 2020. Ecological Diversity in the Terminal Ediacaran Gaojiashan Biota. Earth Science Frontiers, 27(6): 28-46 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Jiang, G. Q., Shi, X. Y., Zhang, S. H., et al., 2011. Stratigraphy and Paleogeography of the Ediacaran Doushantuo Formation (Ca. 635-551 Ma) in South China. Gondwana Research, 19(4): 831-849.  https://doi.org/10.1016/j.gr.2011.01.006 
						
					 | 
			
| 
					 Li, H. K., Zhang, C. L., Xiang, Z. Q., et al., 2013. Zircon and Baddeleyite U-Pb Geochronology of the Shennongjia Group in the Yangtze Craton and Its Tectonic Significance. Acta Petrologica Sinica, 29(2): 673-697 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Liu, P. J., Yin, C. Y., Chen, S. M., et al., 2013. The Biostratigraphic Succession of Acanthomorphic Acritarchs of the Ediacaran Doushantuo Formation in the Yangtze Gorges Area, South China and Its Biostratigraphic Correlation with Australia. Precambrian Research, 225: 29-43.  https://doi.org/10.1016/j.precamres.2011.07.009 
						
					 | 
			
| 
					 Qiu, X. F., Ling, W. L., Liu, X. M., et al., 2011. Recognition of Grenvillian Volcanic Suite in the Shennongjia Region and Its Tectonic Significance for the South China Craton. Precambrian Research, 191(3-4): 101-119.  https://doi.org/10.1016/j.precamres.2011.09.011 
						
					 | 
			
| 
					 Tang, F., Yin, C. Y., Liu, P. J., et al., 2007. Discovery of Diverse Macrofossil Assemblages from the Jiucheng Member of Uppermost Ediacaran in Eastern Yunnan. Journal of Palaeogeography, 9(5): 533-540 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Tang, F., Yin, C. Y., Liu, P. J., et al., 2008. Morphological Comparison of the Ediacaran Miaohe Biota from South China: Implications for Their Affinities and Ecology. Acta Geologica Sinica, 82(5): 601-611, 725-727 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Wang, W., Cawood, P. A., Pandit, M. K., et al., 2021. Fragmentation of South China from Greater India during the Rodinia-Gondwana Transition. Geology, 49(2): 228-232.  https://doi.org/10.1130/g48308.1 
						
					 | 
			
| 
					 Wang, W., Guan, C., Zhou, C., et al., 2014. Exceptional Preservation of Macrofossils from the Ediacaran Lantian and Miaohe Biotas, South China. Palaios, 29(3): 129-136.  https://doi.org/10.2110/palo.2013.085 
						
					 | 
			
| 
					 Wang, Y., 2020. The Ediacaran Wenghui Biota in Northeast Guizhou, South China and the Environment before Its "Explosive Prosperity" (Dissertation). China University of Geosciences, Beijing (in Chinese with English abstract). 
						
					 | 
			
| 
					 Wang, Y., 2009. Megascopic Biota from the Doushantuoian of the Neoproterozoic (Ediacaran) in Guizhou and Zhongqing, South China (Dissertation). China University of Geosciences, Beijing (in Chinese with English abstract). 
						
					 | 
			
| 
					 Wang, Y., Wang, X. L., 2011. New Observations on Cucullus Steiner from the Neoproterozoic Doushantuo Formation of Guizhou, South China. Lethaia, 44(3): 275-286.  https://doi.org/10.1111/j.1502-3931.2010.00241.x 
						
					 | 
			
| 
					 Wang, Y., Wang, X. L., Huang, Y. M., 2007. Macroscopic Algae from the Ediacaran Doushantuo Formation in Northeast Guizhou, South China. Earth Science, 32(6): 828-844 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Wang, Y., Wang, Y., Du, W., 2016. The Long-Ranging Macroalga Grypania spiralis from the Ediacaran Doushantuo Formation, Guizhou, South China. Alcheringa: an Australasian Journal of Palaeontology, 40(3): 303-312.  https://doi.org/10.1080/03115518.2016.1127725 
						
					 | 
			
| 
					 Xiao, S. H., Chen, Z., Pang, K., et al., 2021. The Shibantan Lagerstätte: Insights into the Proterozoic-Phanerozoic Transition. Journal of the Geological Society, 178(1): jgs2020-jgs2135.  https://doi.org/10.1144/jgs2020-135 
						
					 | 
			
| 
					 Xiao, S. H., Muscente, A. D., Chen, L., et al., 2014. The Weng'an Biota and the Ediacaran Radiation of Multicellular Eukaryotes. National Science Review, 1(4): 498-520.  https://doi.org/10.1093/nsr/nwu061 
						
					 | 
			
| 
					 Xiao, S. H., Yuan, X. L., Steiner, M., et al., 2002. Macroscopic Carbonaceous Compressions in a Terminal Proterozoic Shale: A Systematic Reassessment of the Miaohe Biota, South China. Journal of Paleontology, 76(2): 347-376. https://doi.org/10.1666/0022-3360(2002)0760347: mcciat>2.0.co;2 doi:  10.1666/0022-3360(2002)0760347:mcciat>2.0.co;2 
						
					 | 
			
| 
					 Yang, C., Rooney, A. D., Condon, D. J., et al., 2021. The Tempo of Ediacaran Evolution. Science Advances, 7(45): eabi9643.  https://doi.org/10.1126/sciadv.abi9643 
						
					 | 
			
| 
					 Ye, Q., Li, J. Q., Tong, J. N., et al., 2022. A Microfossil Assemblage from the Ediacaran Doushantuo Formation in the Shennongjia Area (Hubei Province, South China): Filling Critical Paleoenvironmental and Biostratigraphic Gaps. Precambrian Research, 377: 106691.  https://doi.org/10.1016/j.precamres.2022.106691 
						
					 | 
			
| 
					 Ye, Q., Tong, J. N., An, Z. H., et al., 2019. A Systematic Description of New Macrofossil Material from the Upper Ediacaran Miaohe Member in South China. Journal of Systematic Palaeontology, 17(3): 183-238.  https://doi.org/10.1080/14772019.2017.1404499 
						
					 | 
			
| 
					 Yuan, X. L., Chen, Z., Xiao, S. H., et al., 2011. An Early Ediacaran Assemblage of Macroscopic and Morphologically Differentiated Eukaryotes. Nature, 470(7334): 390-393.  https://doi.org/10.1038/nature09810 
						
					 | 
			
| 
					 Yuan, X. L., Li, J., Cao, R. J., 1999. A Diverse Metaphyte Assemblage from the Neoproterozoic Black Shales of South China. Lethaia, 32(2): 143-155.  https://doi.org/10.1111/j.1502-3931.1999.tb00533.x 
						
					 | 
			
| 
					 Yuan, X. L., Xiao, S. H., Yin, L. M., et al., 2002. Doushantuo Fossils: Life on the Eve of Animal Radiation. University of Science and Technology of China Press, Hefei (in Chinese with English abstract). 
						
					 | 
			
| 
					 Zhao, Y. L., Chen, M. E., Peng, J., et al., 2004. Discovery of a Miaohe-Type Biota from the Neoproterozoic Doushantuo Formation in Jiangkou County, Guizhou Province, China. Chinese Science Bulletin, 49(20): 2224-2226.  https://doi.org/10.1007/BF03185792 
						
					 | 
			
| 
					 Zhou, C. M., Ouyang, Q., Wang, W., et al., 2021. Lithostratigraphic Subdivision and Correlation of the Ediacaran in China. Journal of Stratigraphy, 45(3): 211-222 (in Chinese with English abstract). 
						
					 | 
			
| 
					 Zhou, C. M., Xie, G. W., McFadden, K., et al., 2007. The Diversification and Extinction of Doushantuo-Pertatataka Acritarchs in South China: Causes and Biostratigraphic Significance. Geological Journal, 42(3-4): 229-262.  https://doi.org/10.1002/gj.1062 
						
					 | 
			
| 
					 Zhu, M. Y., 2010. The Origin and Cambrian Explosion of Animals: Fossil Evidences from China. Acta Palaeontologica Sinica, 49(3): 269-287 (in Chinese with English abstract). 
						
					 | 
			
| 
					 安志辉, 叶琴, 胡军, 等, 2023. 神农架成冰纪宋洛生物群的赋存层位厘定. 地球科学, 48(10): 3799-3811. doi:  10.3799/dqkx.2021.212 
					
					 | 
			
| 
					 安志辉, 叶琴, 童金南, 等, 2024. 鄂西黄陵背斜周缘埃迪卡拉纪灯影组地层序列及其区域地层对比意义. 地球科学, 49(8): 2992-3009. doi:  10.3799/dqkx.2023.041 
					
					 | 
			
| 
					 陈孟莪, 萧宗正, 袁训来, 1994. 晚震旦世的特种生物群落: 庙河生物群新知. 古生物学报, 33(4): 391-403. 
					
					 | 
			
| 
					 陈孝红, 汪啸风, 2002. 湘西震旦纪武陵山生物群的化石形态学特征和归属. 地质通报, 21(10): 638-645 
					
					 | 
			
| 
					 陈孝红, 汪啸风, 王传尚, 等, 1999. 湘西震旦系留茶坡组炭质宏化石初步研究. 华南地质与矿产, 15(2): 15-30. 
					
					 | 
			
| 
					 丁莲芳, 李勇, 胡夏嵩, 等, 1996. 震旦纪庙河生物群. 北京: 地质出版社. 
					
					 | 
			
| 
					 谷昊东, 胡军, 安志辉, 等, 2021. 神农架陡山沱组沉积特征及其对"鄂西海槽"的启示. 地球科学, 46(8): 2958-2972. https://d.wanfangdata.com.cn/periodical/dqkx202108019 
					
					 | 
			
| 
					 华洪, 蔡耀平, 闵筱, 等, 2020. 新元古代末期高家山生物群的生态多样性. 地学前缘, 27(6): 28-46. 
					
					 | 
			
| 
					 李怀坤, 张传林, 相振群, 等, 2013. 扬子克拉通神农架群锆石和斜锆石U-Pb年代学及其构造意义. 岩石学报, 29(2): 673-697. 
					
					 | 
			
| 
					 唐烽, 尹崇玉, 刘鹏举, 等, 2007. 滇东埃迪卡拉(震旦)系顶部旧城段多样宏体化石群的发现. 古地理学报, 9(5): 533-540. 
					
					 | 
			
| 
					 唐烽, 尹崇玉, 刘鹏举, 等, 2008. 华南伊迪卡拉纪"庙河生物群"的属性分析. 地质学报, 82(5): 601-611, 725-727. 
					
					 | 
			
| 
					 王野, 2020. 贵州伊迪卡拉纪瓮会生物群及其"爆发式"繁盛前环境(博士学位论文). 北京: 中国地质大学. 
					
					 | 
			
| 
					 王约, 2009. 黔-渝地区新元古代伊迪卡拉纪陡山沱期宏体生物群(博士学位论文). 北京: 中国地质大学. 
					
					 | 
			
| 
					 王约, 王训练, 黄禹铭, 2007. 黔东北伊迪卡拉纪陡山沱组的宏体藻类. 地球科学, 32(6): 828-844. doi:  10.3321/j.issn:1000-2383.2007.06.014 
					
					 | 
			
| 
					 袁训来, 肖书海, 尹磊明, 等, 2002. 陡山沱期生物群: 早期动物辐射前夕的生命. 合肥: 中国科学技术大学出版社. 
					
					 | 
			
| 
					 周传明, 欧阳晴, 王伟, 等, 2021. 中国埃迪卡拉纪岩石地层划分和对比. 地层学杂志, 45(3): 211-222. 
					
					 | 
			
| 
					 朱茂炎, 2010. 动物的起源和寒武纪大爆发: 来自中国的化石证据. 古生物学报, 49(3): 269-287. 
					
					 |